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Deciduous forests : ウィキペディア英語版
Deciduous

''Deciduous'' means "falling off at maturity" or "tending to fall off", and it is typically used in order to refer to trees or shrubs that lose their leaves seasonally (most commonly during autumn) and to the shedding of other plant structures such as petals after flowering or fruit when ripe. In a more general sense, ''deciduous'' means "the dropping of a part that is no longer needed" or "falling away after its purpose is finished". In plants it is the result of natural processes. "Deciduous" has a similar meaning when referring to animal parts, such as deciduous antlers in deer or deciduous teeth, also known as baby teeth, in some mammals (including humans).
==Botany==
In botany and horticulture, deciduous plants, including trees, shrubs and herbaceous perennials, are those that lose all of their leaves for part of the year.〔(【引用サイトリンク】 title =Trees that lose their leaves )〕 This process is called abscission. In some cases leaf loss coincides with winter — namely in temperate or polar climates.〔(【引用サイトリンク】 title =Science Reference: Deciduous )〕 In other parts of the world, including tropical, subtropical, and arid regions, plants lose their leaves during the dry season or other seasons, depending on variations in rainfall.
The converse of deciduous is evergreen, where foliage is shed on a different schedule from deciduous trees, therefore appearing to remain green year round. Plants that are intermediate may be called semi-deciduous; they lose old foliage as new growth begins.〔(【引用サイトリンク】title=The Illinois - North Carolina Collaborative Environment for Botanical Resources: Openkey Project. Glossary of Botanical Terms. Page 22. )〕 Other plants are semi-evergreen and lose their leaves before the next growing season, retaining some during winter or dry periods.〔Weber, William. 2001. ''African rain forest ecology and conservation an interdisciplinary perspective''. New Haven: Yale University Press. page 15.〕 Some trees, including a few species of oak, have desiccated leaves that remain on the tree through winter; these dry persistent leaves are called marcescent leaves and are dropped in the spring as new growth begins.
Many deciduous plants flower during the period when they are leafless, as this increases the effectiveness of pollination. The absence of leaves improves wind transmission of pollen for wind-pollinated plants and increases the visibility of the flowers to insects in insect-pollinated plants. This strategy is not without risks, as the flowers can be damaged by frost or, in dry season regions, result in water stress on the plant. Nevertheless, there is much less branch and trunk breakage from glaze ice storms when leafless, and plants can reduce water loss due to the reduction in availability of liquid water during cold winter days.
Leaf drop or abscission involves complex physiological signals and changes within plants. The process of photosynthesis steadily degrades the supply of chlorophylls in foliage; plants normally replenish chlorophylls during the summer months. When autumn arrives and the days are shorter or when plants are drought-stressed, deciduous trees decrease chlorophyll pigment production, allowing other pigments present in the leaf to become apparent, resulting in non-green colored foliage. The brightest leaf colors are produced when days grow short and nights are cool, but remain above freezing. These other pigments include carotenoids that are yellow, brown, and orange. Anthocyanin pigments produce red and purple colors, though they are not always present in the leaves. Rather, they are produced in the foliage in late summer, when sugars are trapped in the leaves after the process of abscission begins. Parts of the world that have showy displays of bright autumn colors are limited to locations where days become short and nights are cool. In other parts of the world, the leaves of deciduous trees simply fall off without turning the bright colors produced from the accumulation of anthocyanin pigments.
The beginnings of leaf drop starts when an abscission layer is formed between the leaf petiole and the stem. This layer is formed in the spring during active new growth of the leaf; it consists of layers of cells that can separate from each other.
The cells are sensitive to a plant hormone called auxin that is produced by the leaf and other parts of the plant. When auxin coming from the leaf is produced at a rate consistent with that from the body of the plant, the cells of the abscission layer remain connected; in autumn, or when under stress, the auxin flow from the leaf decreases or stops, triggering cellular elongation within the abscission layer. The elongation of these cells break the connection between the different cell layers, allowing the leaf to break away from the plant. It also forms a layer that seals the break, so the plant does not lose sap.
A number of deciduous plants remove nitrogen and carbon from the foliage before they are shed and store them in the form of proteins in the vacuoles of parenchyma cells in the roots and the inner bark. In the spring, these proteins are used as a nitrogen source during the growth of new leaves or flowers.
==Function==

Plants with deciduous foliage have advantages and disadvantages compared to plants with evergreen foliage. Since deciduous plants lose their leaves to conserve water or to better survive winter weather conditions, they must regrow new foliage during the next suitable growing season; this uses resources which evergreens do not need to expend. Evergreens suffer greater water loss during the winter and they also can experience greater predation pressure, especially when small. Losing leaves in winter may reduce damage from insects; repairing leaves and keeping them functional may be more costly than just losing and regrowing them. Removing leaves also reduces cavitation which can damage xylem vessels in plants. This then allows deciduous plants to have xylem vessels with larger diameters and therefore a greater rate of transpiration (and hence CO2 uptake as this occurs when stomata are open) during the summer growth period.

抄文引用元・出典: フリー百科事典『 ウィキペディア(Wikipedia)
ウィキペディアで「Deciduous」の詳細全文を読む



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